Acts downstream of smo as an intracellular repressor of hedgehog signaling pathway; mainly through the suppression of gli1 activity. This negative regulatory effect is enhanced in conjunction with the suppressor of fused (sufu) protein. Positively regulates gli2a activity by promoting its dissociation from sufu. Involved in the regulation of microtubular dynamics. {ECO:0000269|PubMed:15647323; ECO:0000269|PubMed:24339784}.