Required for oocyte nuclear envelope breakdown before entry of oocyte into spermatheca. In meiotic cells; required for spindle dynamics and probably for spindle attachment to the chromosomes. Zygotic role in the development of the germline and nerve cord. In mitotic cells; plays a role in spindle organization and centrosome maturation. Involved in asymmetric nuclear localization of cdc-25.1 during embryogenesis which affects cell division timing. {ECO:0000269|PubMed:10660671; ECO:0000269|PubMed:18199581; ECO:0000269|PubMed:18305005; ECO:0000269|PubMed:18316412; ECO:0000269|PubMed:20823068}.