Proposed to be part of the KMN network incorporating the MIS12 complex and NDC80 complex that is required for establishing the kinetochore-microtubule interface that aids in chromosome segregation during meiotic and mitotic cell division. Appears to function downstream of hcp-3 (CENP-A) and hcp-4 (CENP-C) in the kinetochore assembly hierarchy. Has a role in the correct localization of spdl-1 and the RZZ complex that is composed of rod-1; czw-1 and zwl-1. {ECO:0000269|PubMed:14522947; ECO:0000269|PubMed:15371340; ECO:0000269|PubMed:15767665; ECO:0000269|PubMed:16273096; ECO:0000269|PubMed:17129783; ECO:0000269|PubMed:18070910; ECO:0000269|PubMed:18765790}.