Serine/threonine-protein kinase which mediates both meiotic and mitotic chromosome segregation. Required for histone H3 'Ser-10' phosphorylation. Phosphorylates mei-S332 within residues 124-126 and stabilizes its association with centromeres during meiosis. {ECO:0000269|PubMed:11266459; ECO:0000269|PubMed:11352945; ECO:0000269|PubMed:16824953}.